In the model, a terrestrial ecosystem is represented as seven interacting components -- plants (divided into leaves, branches, stems, and roots), litter lying on the ground, humus, and stable humus charcoal. The amount of carbon stored in each of these components is a state variable, and is initialized at 0 at the beginning of the simulation.
The seven state variables are linked by a continuous circulation of carbon. Carbon enters the system as the gas carbon dioxide through the plant leaves, and is incorporated into the plants (leaves, branches, stems, and roots) as primary production. Some of the primary production is burnt up in respiration; the remainder constitutes the net primary production of the ecosystem. The model is driven by the net primary production, which may be partitioned among the four components of the plants: leaves, branches, roots, and stems. (A partitioning factor for each of these components for each of the six ecosystems is found as a table at the end of this reading.) You should notice that the four partition coefficients add up to a value of 1.0. For each of the four plant components, carbon uptake is calculated by multiplying the amount of carbon available by its partitioning coefficient. Since the partition coefficient adds up to 1.0, all of the available carbon (for example 27.8 Gt. C/year) should be taken up by the plants.
Carbon moves through the system. Once carbon is taken up by one of the four plant components, it is transformed and transmitted to the other components of the model -- litter on the ground, which in turn is transformed to humus, which is in turn transformed to stable humus charcoal. Note one exception to this; roots are transformed directly into humus, and do not contribute to the litter on the ground. We can categorize each of the processes according to the following table:
Component | Transforms to: | Process name |
Leaves | litter | leaf fall |
Branches | litter | timber fall |
Stems | litter | timber fall |
Roots | humus | sloughage (humification) |
Litter | humus |   |
Humus | stable humus charcoal carbonization |   |
For each transmission or transformation, a "flux" number reflects the fraction of carbon that is passed from one state variable to another. Each of the fluxes for these processes is mathematically defined by multiplying the current value of the state variable by a fractional "transfer coefficient", also found on the data table. For example, the process of transformation from leaves to litter is defined as follows:
Note from the table that the leaves to litter coefficient for most of the ecosystems is 1.0, which implies that all of the carbon found in leaves is fluxed to the state variable of litter.
For the processes of humification and carbonization, values labeled "transfer factors" are shown on the data table. These values are used in a similar manner to the transfer coefficients. They are separated on the data table due to differences in units.
In order to make your model look uncluttered, you will need to design your model carefully.
Extra challenge: how might you develop this model so that the user can change ONE variable to run the simulation for each of the two different ecosystem types? As a hint, you do not necessarily have to use an "if...then" construct to implement this improvement.
  | Units | Tropical Forest Ecosystem 1 | Temperate Forest Ecosystem 2 |
Carbon Dioxide Production | Gt C/yr | 27.8 | 8.7 |
Partition Coefficients | fraction |   |   |
Leaves |   | 0.3 | 0.3 |
Branches |   | 0.2 | 0.2 |
Stems |   | 0.3 | 0.3 |
Roots |   | 0.2 | 0.2 |
Transfer Coefficients (rate constants) | per year |   |   |
Leaves to Litter |   | 1 | 0.5 |
Branches to Litter |   | 0.1 | 0.1 |
Stems to Litter |   | 0.033 | 0.0166 |
Roots to Humus |   | 0.1 | 0.1 |
Litter to Humus |   | 1 | 0.5 |
Humus to Charcoal |   | 0.1 | 0.02 |
Charcoal to Environment |   | 0.002 | 0.002 |
Transfer Factors | fraction |   |   |
Hummification Factors |   | 0.4 | 0.6 |
Carbonization factor for |   | 0.05 | 0.05 |
Humus Decomposition |   | 0.005 | 0.005 |
Other Values | fraction |   |   |
Humus Respiration |   | 0.995 | 0.995 |
Litter Respiration |   | 0.6 | 0.6 |
Stable Humus Respiration |   | 0.002 | 0.002 |